First time posting here. Looking for feedback on mistakes in this hand either from villain or hero.
Live 2-5, loose game
Hero OTB with 53 spades. Stack $750
Villain UTG w QJ spades. Covers hero. Unknown player.

UTG opens for $15. Four callers including hero. Blinds fold
[ Normally 53s is in folding range, but decided to call because A) very multiway B) hi implied odds as table was pretty loose C) absolute IP ]

Flop: 2c 4h 8s (Pot ~$80)
Villain bets $50, folds around. Hero calls.

Turn: 7s (Pot ~180)
Villain bets $125. Hero calls

River: Ad (Pot ~430)
Villain shoves, hero calls

Potential mistakes:
1. Hero calling with 53s?
2. Villain bluffing on flop into 4 players OOP?
3. Hero turn call? (absolutely did not expect villain to have spades. Given bet into 4 players on flop, range looks almost exclusively like an overpair or set, which supports implied odds on river if the OESD+FD comes in. Maybe could see AKs playing out this way -but don’t even see this supporting the Flop bet given the field. )
4. Villain shove on river?
5. Other?

Thank you