fold23bb

Hi, I am practicing my range reading by looking at positional ranges and bet sizing. Came across this situation (I wasn't involved), but tried to range the players (and failed miserably).

6 max (.25/.5 NL)
Villain 1 (MP) 55 hands - VPIP 39, PFR 24, 3Bet 10.3%
Villain 2 (CO) 54 hands - VPIP 23, PFR 11, 3Bet 0%

Preflop:
Villain 1 (MP): Raises 1.5$
Villain 2 (CO): Calls 1.5$
Villain 3 (BB): Calls 1$
Pot Size: 4.75$

Flop: 8s5c2c
Villain 3 (BB): Checks
Villain 1 (MP): Checks
Villain 2 (CO): Bets 3.13$
Villain 3 (BB): folds
Villain 1 (MP): Calls 3.13
Pot Size: 11.01$

Turn: 8s5c2cKs
Villain 1 (MP): Checks
Villain 2 (CO): Bets 5.50$
Villain 1 (MP): Raise 13.75$
Villain 2 (CO): Calls 8.25$
Pot Size: 38.51$

River: 8s5c2cKs7c
Villain 1 (MP): Bets 19.25$
Villain 2 (CO); Calls 19.25$

After using Flopzilla to range them, I came to the conclusion of both having club Flush, with my estimate being that Villain 1 probably had KcXc (which is why he bet 80% pot on Turn, had top pair and a flush draw of clubs). Neither have Ac nut flush.

Actual Hands: Villain 1 (MP): AdQs and Villain 2 (CO) AcKh

4 questions here:

1. Where (is it likely) did I go completely wrong in my ranging (probably not paying attention to CO's PFR and 3bet stats)?
2. Why was Villain 1 bluffing with AdQs? I don't have any GTO solvers so I don't know how this should be looked at.
3. How should Villain 2's bet sizing on flop and turn be interpreted? Villain 2 (with Ac) was blocking a lot of club combos in MP's range, but apart from that, what was he really going for? Was that a 'scare off' bet on the Flop with nothing made, and a value bet on the Turn (with the K pair), but just a call on the check-raise on account of the flush possibilities? This I can rationalize now that I know his cards, but without knowing them how should Villain 2's bet sizing be interpreted?
4. What should Villain 1 have bet on river to pull of a successful bluff? Clearly a half pot raise didn't do the trick.

Any guidance would be much appreciated.